The settlements were an invasion of their land. The settlers took food, space, and resources that they had no natural right to.
The product of exposure, life experiences and learning of an individual
Providing harsh criticism to other players is a natural part of being a good leader - this is true. The reason for this is that good leaders will often be able to accept a harsh critique coming their way and also be able to give a harsh critique which a player deserves.
DNA that makes them grow better, bigger, and faster