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Answer:
The last one is the answer
Important ideas of life or human nature expressed
The only option that doesn't really fit here is "ECONOMICS".
Darwin's theory of evolution opposed the ideas of the church that god was responsible for all progress and the being of humans. This is clearly a challenge to religion.
Until Darwin made his discoveries, science had been unable to truly understand the idea of evolution and for that reason he was also challenging the scientific world with a revolutionary theory.
Education is a mix of the two. At the time, education was highly linked to the church and would have shared morals along with beliefs. Science was clearly being taught in all schools at the time and of course Darwin's ideas ran contrary to the ideas that were being disseminated at the time.