I think the organisms are compared to each other.
The guppies evolved at a rate between 3700 and 45,000 Darwin's. For comparison, artificial-selection experiments on mice show rates of up to 200,000 Darwin's while most rates measured in the fossil record are only 0.1 to 1.0 Darwin.
Guppies introduced to a low-predation environment adapted to their new environment by producing larger and fewer offspring with each reproductive cycle. No such adaptation was seen in guppies that colonized a high-predation
With each generation, more guppies passed on the gene for small spots to their offspring. The rocks became smaller. More guppies with small spots survived long enough to pass on the gene for small spots to their offspring the rocks became smaller.
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