I can't read the paper, so all I can say is read Darwin's Survival of the Fittest and you will understand evolution.
The answer is ‘in the dermal papillae of hairless skin’. Dermal papillae is the layer of the skin (upper dermis) with numerous blood vessels and nerves that serve the epidermis of the skin. Tactile corpuscles (nerves) are mostly found in this region and occur mostly in hairless thick skin such as in finger and toe tips.
Not sure but i think cohesion forces
4-Growth, maintenance, balance fluids, maintain proper hp
2-Actin, collagen
