When a plant has been without water for a long time, the central vacuoles lose water, the cells lose shape, and the whole leaf wilts. Plants often store sugars, ions, some proteins and occasionally pigments inside the vacuole.
It contributed to the rise of the Ottoman Empire. When the Byzantine Empire ended, the Ottomans received a strong trade port affecting Mediterranean-black sea trade zones.<span />
By implying that blacks and whites are 'separate but equal'.
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