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Some mutations have a positive effect on the organism in which they occur. They are called beneficial mutations. ... They increase an organism's changes of surviving or reproducing, so they are likely to become more common over time. There are several well-known examples of beneficial mutations.
The horse and donkey, though closely related by sharing a common ancestor, are different species. They have different number of chromosomes hence pairing of homologous chromosomes during fusion of gametes becomes complicated. The mule will have an extra chromosome from the horse hence will have abnormalities such as sterility. A Mule is unable to reproduce due to this same phenomenon. Homologous chromosomes are not well paired for meiosis I.
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