Answer: The size.
Explanation: On the one hand the Emperor Penguin is bigger reaching up to 48 Inches (120cm) of height and 100 Lb in weight, on the other hand the Galápagos Penguin is much smaller. For example a large adult usually goes up to 21 Inches (53cm) and 5 Lb of weight.
This is directly related to the temperature of the environment in which each one develops, because a body exchanges heat in two dimensions through the surface. So, it's easier for a big creature to maintain core temperature stable.
This is really important in environments like the Arctic where temperature goes below zero and the opposite happen in the Equator where there is a tropical weather all the time.
According to sources, the most probable answer to this query is allostasis. This the body's way of being able to cope with stressful situations and respond accordingly.
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