The four organelles that contribute to the synthesis, transport, packaging, and export of proteins to the extracellular environment are Nucleus, RIBOSOMES, rough endoplasmic recticulum and Golgi apparathus.
Explanation:
The nucleus is the organelle that produced RIBOSOMES.
RIBOSOMES are organelles responsible for protein synthesis production of proteins.
The protein produced by the RIBOSOMES are then gathered in the rough endoplasmic recticulum . It then export and transport the protein to Golgi apparatus. The Golgi apparatus modifies and package the protein.
