"The natural predators and prey of arachnids in tropical ecosystems"
Explanation:
Because it's the only one that simply says 'arachnids' and not 'brown widow spiders', and instead of specifically saying 'Natural predators' it says 'natural predators and prey'
1 "The natural predators and prey of arachnids in tropical ecosystems".
Explanation
I believe this is right because while the other anwsers bring up actual spider 1 just says arachnids which is simply a class of joint-legged invertebrate animals. they also include predators and prey.