They held desired resources which included land and labor. Hope this helps.
The feature that helped Timbuktu make it a center of intellectual advances and culture is the "Library." Books themselves are very intellectual in its nature. It was in the <span>Askia dynasty that Timbuktu felt its prosperity in intellectual advances. </span>
According to a source, osmoregulation is primarily accompanied by sodium and potassium pumps that works together with regulating the osmotic pressure within these organisms and fishes.
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