Hamilton's rule, mathematical formula that supports the notion that natural selection favors genetic success, not reproductive success per sea. But by the recipient of the altruism, ℂ is the cost (in number of offspring equivalents) suffered. The altruistic behaviour, and r is the genetic relatedness of the altruist to the beneficiary.
Explanation:
I am not fursure on what your answer would be but I hope that what I put can help you find your answer.
