During Meiosis 1 in Prophase 1 crossing over occurs. The two chromosomes switch part of their genetic information. Also independent assortment occurs during meiosis which is the formation of random combinations of chromosomes in meiosis and of genes on different pairs of homologous chromosomes.
It is with sperm and eggs, which are housed in spores in both species, so I’d say spores is the safest answer.
