He was a close friend to Charles Darwin so he helped him out in creating the theory of evolution so I would say C: He developed the idea of inheritance of acquired characteristics.
In DNA methylation, the most frequently methylated nitrogenous base is the cytosine. Its methylation is widespread in prokaryotes and eukaryotes. Its rate of DNA methylation differs greatly between different species like Arabidopsis thaliana which is 8%, Physarum, 4%, Mus musculus, 2.3%, and etc.