Leonardo Da Vinci is quite the man, he had popular interests not only in art but in science, engineering, mathematics, sculpture, architecture, and anatomy. That’s just to name a few. He popularly made artworks like the Mona Lisa, the last supper, the Vitruvian man, Saint John the Baptist, Salvator mundi and much more. High renaissance / Italian Renaissance era.
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I believe the correct answer is: human beings (individuals).
The artistic epoch of Humanism emphasizes the significance
of rational thought, rather than adherence to the principles of faith. Therefore,
the influence of the Humanism on Renaissance art was a focus on dignity,
freedom and the human potential of realizing self-actualization. But, even though
individual is in the focus, the God was not abandoned.