The last one. The only scenario where you wouldn't put a comma in the first quote is if it's a completed remark that uses particular punctuation such as a quotation or exclamation mark, but never a period. Hope this helped.
Answer:
Helpful or Harmful?
Explanation:
— Herbicide-tolerant genetically modified (GM) crops have led to an increase ... food costs, they also help to combat hunger and related problems in various ... amounts of these pesticides are unclear, but studies have linked them to a ... Glyphosate is the most widely used herbicide in the world by volume [9].
He should have a good hook in place and then have a good connective bridge before writing the thesis on the paper.
I hope this helps.