In broad terms, finches. In specific terms, he observed how over time, the beak sizes and shapes of said finches changed to adapt to the food sources available.
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Answer:
atmospheric pressure
Explanation:
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There is only one measure of "evolutionary success": having more offspring. A "useful" trait gets conserved and propagated by the simple virtue of there being more next-generation individuals carrying it and particular genetic feature "encoding" it. That's all there is to it.
One can view this as genes "wishing" to create phenotypic features that would propagate them (as in "Selfish Gene"), or as competition between individuals, or groups, or populations. But those are all metaphors making it easier to understand the same underlying phenomenon: random change and environmental pressure which makes the carrier more or less successful at reproduction.
You will sometimes hear the term "evolutionary successful species" applied to one that spread out of its original niche, or "evolutionary successful adaptation" for one that spread quickly through population (like us or our lactase persistence mutation), but, again, that's the same thing.
The geosphere has four subsystems called the lithosphere, hydrosphere, cryosphere, and atmosphere. Because these subsystems interact with each other and the biosphere, they work together to influence the climate, trigger geological processes, and affect life all over the Earth.