The answer is macrophages. They either actively invade these leukocytes or are phagocytosed, divide in the cells and cause lysis. The promastigotes that invade these leucocytes are transformed into amastigotes in the macrophages. These amastigotes continue attacking other healthy macrophages while others migrate to the mid gut.
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To quote another answer I seen from this same question "I would say that the theme which is reinforced in this excerpt is impossibility of certainty - Hamlet is uncertain what he should do next, and he expresses that uncertainty in this soliloquy. "
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