Explanation:natural disasters: this affect population where by organisms leave there area or habitat for another area due to unfavorable conditions eg. flood, bush fire, storms, earthquake etc. and can also lead to death.
Emigration:this is when organisms leave an area for another either in search of food, breeding purpose this reduses population where by organisms may never return.
scarcity of food:this occur when there is insufficiency or no food in an area which makes organisms leave their habitat for another in search of food and might even lead to death.
The endoderm layer may have developed abnormally" is the possible reason among the choices given in the question. The correct option among all the options that are given in the question is the third option or the last option. I hope that this is the answer that has actually come to your great help.