The mantle is a layer inside a planetary bodybounded below by a core and above by a crust. Mantles are either made of rock or ices, and are generally the largest and most massive layer of the planetary body. Mantles are characteristic of planetary bodies that have undergone differentiation by density. The mantle is bounded on the bottom by the planetary core and on top by the crust. In addition to the Earth, the other terrestrial planets, a number of asteroids, and moonshave mantles.
A mother passes on her mt(DNA) or mitochondrial DNA which sometimes includes genetic material but a father only passes down a Y chromosome if he has a son. No genetic material.
If the web happened to look like this one I attached then the anwser would be shrubs. If you take a look at the arrows the shrubs energy is moving to rats, insects and lizards which in turn feeds the coyotes. The starting point is shrubs and in general most food webs start with plants.