I would say immigration only.
emigration would theoretically reduce the genetic variation as unique individuals leave, but immigration would bring unique traits into circulation in an ecosystem.
I'm not super great at biogy tho so this might not be right... :)
I do not understand your question fully because there might be some context missing to it:
Having more nuclei is not something caused by the lack of a process or stage. Some muscle cells usually fuse together, which means they become one. But before they became one, each had their own nuclei. And when they fused, each one kept their nuclei, making one cell with more than one nuclei.
Those cells are called multinucleated cells.
Hope it helped,
BioTeacher101
If you want faster replys or even replys at all, you need to give all the info.